From the crystal composition of your D273N mutant of NagA, the side chain of His-143 is three.0 Å faraway from the phosphonate oxygen from the mimic with the tetrahedral intermediate (21). This consequence implies this team could aid the activation with the carbonyl group with the substrate at the side of an conversation Using the metal ion bound inside the Lively web-site.
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You will find there's proton transfer within the sure drinking water molecule to Asp-273 which is followed by nucleophilic assault within the carbonyl carbon and formation from the tetrahedral intermediate. The collapse of your tetrahedral login naga26 intermediate is facilitated by proton transfer from Asp-273 to your leaving group amine of your item, d-glucosamine-six-phosphate. The products depart the Lively internet site and Yet another spherical of catalysis commences With all the charging in the metal Middle having a molecule of water through the solvent.
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